Research Fragments

A scientific blog by Guilhem Doulcier.

1st June 2020

Probability and fitness


This is an interactive version of Figure 3 and Figure 4 of the article Taming Fitness: Organism-Environment Interdependencies Preclude Long-Term Fitness Forecasting (Doulcier, Takacs and Bourrat, 2021).

dice
red flip
blue flip
red heads
red tails
blue heads
blue tails

Let :

The die is cast before each coin toss: No irreversible changes

This corresponds to Section 4, Game 1 in the article.
Rules of game 1

The average-over-all-possible coins score for the heads player is 0 (0 for the red coin and 0 for the blue coin. E = (0 + 0)/2 = 0).

Independant rounds variant

The properly weighted average-over-all-possible coins score for the heads player is 0 (0 × 0 for the red coin and 0 × 0 for the blue coin. E = (0×0 + 0×0) = 0).

Both the average score per round Yn and the average score per outcome Zk converge toward the properly weighted average-over-all-possible coins score.

Markovian rounds variant

The properly weighted average-over-all-possible coins score for the heads player is 0 (0 × 0 for the red coin and 0 × 0 for the blue coin. E = (0×0 + 0×0) = 0). [The weights are computed using the eigenvalues of the transition matrix.]

Both the average score per round Yn, and the average score per outcome Yk converge toward the same value for the Tails player. (The convergence may only become visible for large sample size, but can be analytically proven using Markov-chains theory).

The die is cast once at the beginning: Irreversible changes

This corresponds to Section 4, Game 2 in the article.
Rules of game 2

Even though both red and blue coins are "possible" the average score per round Yn never converges toward the average-over-all-possible-coins score. This is because once the die has been cast, the colour of the coin never changes. Note that the average score per outcome Zk still converges toward the properly weighted average-over-all-possible-coins score of for the heads player.

Rare events change the environement and rare events change the side of a player: Mutations

This corresponds to Section 5 in the article.
Mutations are marked as * in the outcome.

Note that both the average score per round Yn, and the average score per outcome Zk converge toward the same value for both players. (The convergence may only become visible for large sample size, but can be analytically proven using Markov-chains theory).

References

Source: main.js Software used: d3js.

The article:

Taming fitness: Organism-Environment Interdependencies Preclude Long-Term Fitness Forecasting. Guilhem Doulcier, Peter Takacs, Pierrick Bourrat, BioEssays, 2020.